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Greater Short-toed Lark, Calandrella brachydactyla

The Greater Short-toed Lark (Calandrella brachydactyla) is a small passerine bird. The current scientific name is from Ancient Greek. The genus name, Calandrella, is a diminutive of kalandros, the calandra lark, and brachydactila is from brakhus, "short", and daktulos, "toe".

It breeds in southern Europe, north-west Africa, and across the Palearctic from Turkey and southern Russia to Mongolia. During migration they form large, tight flocks that move in unison; at other times they form loose flocks.

Distribution and habitat
All but some southernmost populations are migratory, wintering south to the southern edge of the Sahara and India. This species is a fairly common wanderer to northern and western Europe in spring and autumn. Populations breeding in the Iberian Peninsula winter south of the Sahara in Africa.

Here they prefer crop land and dry pastures with short shrubs while the syntopic lesser short-toed larks (Calandrella rufescens) prefer drier areas.

This is a common bird of dry open country and cultivation. It nests on the ground, laying two to three eggs. Its food is seeds and insects, the latter especially in the breeding season.

In colonial India, they were hunted for food as ortolans.

They visit parts of South Asia in large flocks during winter and are sometimes attracted to short grass areas along aerodromes and become a bird strike risk to aircraft.

Greater Short-toed Lark, Calandrella brachydactyla
Range map
Range map from - Ornithological Portal is one of those MUST visit pages if you're in to bird watching. You can find just about everything there

Greater Short-toed Lark, Calandrella brachydactyla
Range of C. brachydactyla - Click picture for full size map
By SanoAK: Alexander Kürthy - Made with Natural Earth. Free vetor and raster map data @
Range map from BirdLife International 2018. Calandrella brachydactyla.
The IUCN Red List of Threatened Species 2018: e.T103766207A132042070.
Downloaded on 22 July 2019 as visual indicator of distribution.,
CC BY-SA 4.0,

Taxonomy and systematics
The greater short-toed lark was described by the German naturalist Johann Leisler in 1814 and given the binomial name Alauda brachydactila. This lark is now placed in the genus Calandrella that was established by another German naturalist, Johann Jakob Kaup, in 1829.

The specific name brachydactyla is from the Ancient Greek βραχυδακτυλος brakhudaktulos "short-toed" from brakhus "short" and daktulos "toe". The alternate name short-toed lark may also be used for three other species in the genus Calandrella.

The Mongolian short-toed lark was formerly considered as a subspecies of the greater short-toed lark (as C. b. dukhunensis) until split in 2016 by the IOC. Formerly, some authorities also considered the red-capped lark to be either conspecific (as C. cinerea) with or a subspecies (as C. b. cinerea) of the greater short-toed lark.


Eight subspecies are recognized:

• European greater short-toed lark (C. b. brachydactyla) or Palestine short-toed lark - (Leisler, 1814): Found in southern Europe, on Mediterranean islands and in north-western Africa.
• Hungarian greater short-toed lark (C. b. hungarica) - Horváth, 1956: Found in Hungary and northern Serbia
• North African greater short-toed lark (C. b. rubiginosa) - Fromholz, 1913: Found in northern Africa
• Levant greater short-toed lark (C. b. hermonensis) - Tristram, 1865: Originally described as a separate species. Found from southern Turkey and Syria to north-eastern Egypt
• BIrelandcik greater short-toed lark (C. b. woltersi) - Kumerloeve, 1969: Found in southern Turkey and north-western Syria
• Transcaucasian greater short-toed lark (C. b. artemisiana) - Banjkovski, 1913: Found from central Turkey and Transcaucasia to north-western Iran
• Steppe greater short-toed lark (C. b. longipennis) - (Eversmann, 1848): Originally described as a separate species in the genus Alauda. Also known as Eastern short-toed lark (a name also used by the Asian short-toed lark) or Yarkand short-toed lark. Found from Ukraine and southern Russia to south-central Siberia and southern Mongolia
• C. b. orientalis - Sushkin, 1925: Found in central Siberia, northern Mongolia and northern China

Several subspecies have been named but there is considerable geneflow and the species itself forms part of a larger complex. This is a small pale lark, smaller than the skylark.

It is dark-streaked greyish-brown above, and white below, and has a strong pointed bill that is pinkish with a grey culmen.

It has a pale supercilium, dark patches on each side of its neck and a dark tail. Some birds in the west of the range have a rufous crown. The sexes are similar. The greater short-toed lark is paler than the Mongolian short-toed lark which also has a shorter bill.

In winter they fly in large and compact flocks that swing in synchrony.

Care must be taken to distinguish this species from other similar Calandrella larks, such as the lesser short-toed lark.

The nominate form breeds in Europe (Iberia, France, Italy, the Balkans and Romania) and winters in Africa. Subspecies hungarica breeds in the eastern parts of Europe while rubiginosa breeds in north-western Africa. Subspecies hermonensis (sometimes including woltersi) breeds in Turkey, Syria and Egypt.

Subspecies artemisiana (considered by some to be synonymous with longipennis) breeds in Asia Minor and winters in southern west Asia. Subspecies longipennis breeds in Ukraine, Mongolia and Manchuria and winters in South Asia mainly in the drier zone of north-western India.

The song varies between a dry twittering and a more varied and imitative melody. Flocks will often fly together to water in the mornings at favourite spots. In the evenings they roost in open ground, with each bird squatting in a small depression made in the soil.

Length: 15 cm
Wingspan: 27 to 32 cm
Weight: 18 to 25 g
Longevity: 5 years

Listen to the Greater Short-toed Lark

Remarks from the Recordist

Calls as the bird was on the ground.

Conservation status
Conservation status
Least Concern (IUCN 3.1)
IUCN Red List of Threatened Species. 2012. Retrieved 26 November 2013.

From Wikipedia, the free encyclopedia
From Wikipedia, the free encyclopedia

Sighted: (Date of first photo that I could use) 7 March 2021
Location: eBird hotspot: Episkopi (Επισκοπή) Environmental Centre and Village, Cyprus

Greater Short-toed Lark, Calandrella brachydactyla
Greater Short-toed Lark - 15 March 2021
eBird hotspot: Mandria (Μανδριά), Cyprus

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